Curriculum Vitae
Education
Ph.D. in Marine Ecology, Utrecht University, The Netherlands
2018 – 2023
M.Sc. in Marine Biology, University of Chinese Academy of Sciences, China
2015 – 2018
B.Sc. in Aquaculture Science, Yantai University, China
2011 – 2015
Academic Positions
Postdoctoral Fellow, Linnaeus University, Sweden
2026 – Current
Postdoctoral Researcher, The University of Hong Kong, Hong Kong SAR, China
2023 – 2025
Research Assistant, Royal Netherlands Institute for Sea Research (NIOZ), The Netherlands
2023
Research Focus
- Environmental variability and marine heatwaves
- Benthic invertebrate physiology and behaviour
- Bioturbation and sediment–ecosystem feedbacks
- Variance- and threshold-based approaches to ecological resilience
- Mesocosm experiments and reproducible data workflows (R / Python)
Publications
Zhou, Z., Fivash, G. S., Cozzoli, F., Walles, B., Troost, K., Ysebaert, T., & Bouma, T. J. (2025).
Compound extreme events reshuffle the stacked odds in the gamble between native and introduced bivalves.
Global Ecology and Conservation, e03918.
Lehuen, A., Oulhen, R.-M., Zhou, Z., de Smit, J., van IJzerloo, L., Cozzoli, F., Bouma, T. J., & Orvain, F. (2024).
Multispecies macrozoobenthic seasonal bioturbation effect on sediment erodibility.
Journal of Sea Research, 201, 102525.
Zhou, Z., Grandjean, T. J., de Smit, J., van Belzen, J., Fivash, G. S., Walles, B., Beauchard, O., van Dalen, J., Blok, D. B., van IJzerloo, L., et al. (2024).
Sediment dynamics shape macrofauna mobility traits and abundance on tidal flats.
Limnology and Oceanography, 69(10), 2278–2293.
Zhou, Z., Steiner, N., Fivash, G. S., Cozzoli, F., Blok, D. B., van IJzerloo, L., van Dalen, J., Ysebaert, T., Walles, B., & Bouma, T. J. (2023).
Temporal dynamics of heatwaves are key drivers of sediment mixing by bioturbators.
Limnology and Oceanography.
Zhou, Z. (2023).
Benthic macrofauna under extremes: Unraveling the response strategies from individual behavior to community structure in tidal flats.
PhD Thesis, Utrecht University.
Zhou, Z., Bouma, T. J., Fivash, G. S., Ysebaert, T., van IJzerloo, L., van Dalen, J., van Dam, B., & Walles, B. (2022).
Thermal stress affects bioturbators’ burrowing behavior: A mesocosm experiment on common cockles (Cerastoderma edule).
Science of the Total Environment, 824, 153621.
Chen, L., Lutaenko, K. A., Li, X., Li, X., Zhou, Z., Li, B., Pavlyuk, O. N., & Tarasova, T. S. (2020).
Long-term changes of marine subtidal benthic communities in North East Asia (Yellow and Japan seas) in a global change context: A review.
Aquatic Conservation: Marine and Freshwater Ecosystems, 30(7), 1451–1475.
Li, X., Chen, L., Zhou, Z., Li, B., Liu, X., Yang, L., Liu, B., & Song, B. (2020).
Spatio-temporal variation of subtidal macrobenthic fauna and ecological assessment of Longkou Artificial Island construction in the Bohai Sea, China.
Journal of Oceanology and Limnology, 38(6), 1811–1824.
Zhou, Z., Li, X., Chen, L., Li, B., Wang, C., Guo, J., Shi, P., Yang, L., Liu, B., & Song, B. (2019).
Effects of diesel oil spill on macrobenthic assemblages at the intertidal zone: A mesocosm experiment in situ.
Marine Environmental Research, 152, 104823.
Yang, L., Chen, L., Li, X., Zhou, Z., Liu, B., Song, B., Li, B., & Li, B. (2019).
Effect of seasonal hypoxia on macrobenthic communities in the Muping Marine Ranch, Yantai, China.
Biodiversity Science, 27(2), 200.
Li, B., Bouma, T. J., Wang, Q., Soissons, L. M., Cozzoli, F., Feng, G., Li, X., Zhou, Z., & Chen, L. (2019).
Effects of the mud snail Bullacta exarata on oxygen and nutrient fluxes at the sediment–water interface in the Huanghe River Delta, China.
Acta Oceanologica Sinica, 38(8), 48–55.
Zhou, Z., Li, X., Chen, L., Li, B., Liu, T., Ai, B., Yang, L., Liu, B., & Chen, Q. (2018).
Macrobenthic assemblage characteristics under stressed waters and ecological health assessment using AMBI and M-AMBI: A case study at the Xin’an River Estuary, Yantai, China.
Acta Oceanologica Sinica, 37(5), 77–86.
Li, B., Zhou, Z., Li, B., Wang, Q., Li, X., & Chen, L. (2018).
Size distribution and reproductive cycle of Asterias amurensis (Echinodermata: Asteroidea) in China.
Acta Oceanologica Sinica, 37(6), 96–103.
Chen, L. L., Wang, Q. C., Li, X. J., Zhou, Z., & Li, B. Q. (2016).
Long-term trends of macrobenthos in the southern Bohai Sea, China.
Scientia Sinica Vitae, 46(9), 1121–1134.
Li, X., Zhou, Z., Chen, L., & Li, B. (2016).
Characteristics of macrobenthic communities in the estuary of the Dagujia River and adjacent waters in Yantai, Shandong, China.
Biodiversity Science, 24(2), 157–165.
Li, B., Li, X., Zhou, Z., et al. (2016).
Ecological effects of reclamation on benthic communities.
Guangxi Sciences, 23(4), 293–298.
Li, X., Zhou, Z., Chen, L., Li, B., Liu, T., Ai, B., Yang, L., Liu, B., & Wang, S. (2017).
Effects of coastal reclamation on benthic macrofauna in the Caofeidian coastal area, Bohai Bay, China.
Oceanologia et Limnologia Sinica, 48(3), 617–627.
Li, B., Li, X., Bouma, T. J., Soissons, L. M., Cozzoli, F., Wang, Q., Zhou, Z., & Chen, L. (2017).
Analysis of macrobenthic assemblages and ecological health of the Yellow River Delta using AMBI and M-AMBI.
Marine Pollution Bulletin, 119(2), 23–32.
Conferences & Invited Talks
International Temperate Reefs Symposium (ITRS), 2025
Oral presentation; ITRS CommitteeInternational Symposium on Marine Science & Technology (ISMST), 2025
Oral presentation; Zhejiang Ocean UniversityNWO-Life Conference, 2023
Prompt talk and poster presentation; Dutch Research CouncilNCK Days, 2023
Oral presentation; Netherlands Centre for Coastal ResearchInternational Symposium on Coastal Resources and Environment (CORE), 2021
Oral presentation; International Association of Hydro-Environment Engineering and Research (IAHR)Graduate Forum on Marine Life Sciences & Shandong Provincial Summer School for Marine Life Sciences Graduate Students, 2017
Oral presentation (Chinese); Bureau of Frontier Sciences and Education, Chinese Academy of Sciences & Department of Education of Shandong ProvinceThe 12th National Symposium on Biodiversity Science and Conservation, 2016
Poster presentation; Bureau of Science and Technology for Development, Chinese Academy of Sciences & Biodiversity Committee of the Chinese Academy of SciencesSymposium on Oceanography and Limnology under Global Change: Regional Responses and the Future Ocean, 2016
Poster presentation; Chinese Society for Oceanology and Limnology & Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences
Co-supervision on Internship & Thesis
Cristian Martin, Ghent University (2020.01 – 2020.06)
IMBRSea Joint MSc ThesisSimon Hof, HAN University of Applied Sciences (2020.08 – 2021.01)
Internship Research ProjectMax Wielders, HAS University of Applied Sciences (2021.03 – 2021.06)
Internship Research ProjectJanneke de Bresser, Utrecht University (2021.04 – 2021.07)
BSc ThesisQuinn Bommelé, HZ University of Applied Sciences (2021.06 – 2021.11)
BSc ThesisTony Chi Wa So, The University of Hong Kong (2025.03 – 2025.06)
MSc Thesis
Funding & Grants
Environment and Conservation Fund (Hong Kong)
Assessing impacts of sea urchin grazing on Sargassum biodiversity in Hong Kong waters
(2025, under review)China Scholarship Council (CSC) — Chinese High-Level University Postgraduate Project
(Grant No. 201804910683, Awarded, 2018–2022)
Academic Service
Peer Review for Journals:
- Global Change Biology
- Estuarine, Coastal and Shelf Science
- Wetlands
- Environmental Sciences Europe
- AIMS Geosciences
Awards & Certifications
- Academician Liu Ruiyu Marine Science Scholarship (2018)
- Outstanding Student Leader Award, University of Chinese Academy of Sciences (2018)
- Excellent Report Award, Graduate Forum on Marine Life Sciences, Chinese Academy of Sciences (2017)
- First Place Award, Shandong Provincial Undergraduate Science & Technology English Contest (2014)
- Cambridge Business English Certificate — C1